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C4 Pathway, CO2 (in atmosphere) dissolves in cytoplasm Hydrogencarbonate HCO3-, Hydrogencarbonate HCO3- PEP Carboxylase C4 acids (e.g. malate, aspartate), C4 acids (e.g. malate, aspartate) Decarboxylation C3 acid, C4 acids (e.g. malate, aspartate) Decarboxylation CO2, Plasmodesmata , CO2 Calvin Cycle, Phosphoenolpyruvate PEP Carboxylase C4 acids (e.g. malate, aspartate), , C3 acid Regeneration Phosphoenolpyruvate,