Roughness DNA = <S4
S4
S4
S3
S3
S3
S3
S3
S4
S3
S2
S2
S5
S2
S3
S4
S3
S3
S3
S3
S3
S3
S3
S5
S4
S4
S5
S5
S4
S4
S3
S3
S2
S3
S4
S4
S4
S4
S3
S4
S4
S3
S2
S4
S4
S4
S3
S4
S4
S4
S3
S2
S3
S2
S4
S4
S3
S3
S4
S3
S3
S1
S3
S4
S3
S3
S4
S5
S3
S3
S5
S4
S3
S3
S4
S4
S3
S3
S1
S3
S3
S3
S4
S2
S2
S3
S3
S3
S2
S3
S3
S2
S4
S3
S5
S2
S2
S2
S4
S3
S3
S5
S3
S4
S3
S3
S4
S4
S3
S5
S3
S4
S4
S3
S3
S4
S5
S4
S3
S3
S3
S4
S1
S4
S3
S3
S4
S2
S3
S2
S4
S5
S3
S3
S4
S4
S2
S4
S4
S5
S5
S4
S4
S4
S3
S3
S3
S2
S5
S4
S4
S3
S4
S5
S2
S3
S3
S1
S3
S4
S4
S3
S4
S4
S2
S4
S3
S3
S3
S3
S2
S2
S3
S4
S4
S3
S4
S2
S2
S4
S3
S2
S2
S3
S4
S4
S4
S2
S2
S5
S5
S4
S3
S3
S5
S5
S5
S3
S4
S4
S3
S3
S3
S4
S4
S4
S3
S3
S4
S2
S3
S2
S3
S4
S4
S4
S5
S4
S3
S2
S2
S2
S2
S3
S3
>